The el-509's are basically a drop in replacement for the 6lq6/6me6 or the 6mj6's all are 6.3v filament tubes...
You have to change the socket configuration wiring as the grid and heater pins do differ from the 6lq6 vs the el-509's. The existing sockets are the same pin count and el's will fit. I however preferred to change them as the ceramic replacements I found were a much stouter version than the Bakelite versions D&A used in most of their amps...
Now a little logic...11 tube sockets at $2.50 each +/- = say $28.00
11/ el-509's at best price $15.00 each (more like $20-$22 with shipping)....
11 x $20.00 = $220.00 + sockets $28.00 = $248.00 parts only
Add in 20 or more hours labor gutting the existing RF deck / installing new sockets and wiring...how much is your time worth?
Now you understand why you can buy or carry those "Things" away for a song!
The 4 tuber's like the Mav 250's much less of and investment and yield an amp project for less than parts cost of one of those monster autrosities...IMHO
Will you learn something....most certainly!....but the output difference maybe 1 S unit(6db)...IMHO...no not really....but your call to make
Even if you could find 6lq6's in a lot for $25 each....$300 just to re-tube her as she sits.
Now your question...yes you can read the B+ voltage with the tubes removed...but watch your butt...only with a GOOD meter and test leads rated at a 1000 volts or more...Be careful the power in that amp will kill you in an instant....if it does not...you may wish you did not wake up!
Please be careful
Adding tubes together like 5/24v to equal 120v filament, again IMHO is a death wish or a good reason to have the FDNY on stand-by when you throw the switch...I won't even touch a Firebird or the likes that does not contain a separate filament winding rated for the tubes (6.3/12v OK) and weird combinations like the old TV's used...nope not anymore!
There is many a story there by anyone who worked on those...
GL...Be careful
All the Best
BJ