So, here's a parallel (maybe) experience from the Heathkit SB-220 amplifier.
Not the same thing, but it shares the same basic circuit.
To start, we have to consider that the negative side of the HV supply is not ground. Simpler stuff like sweep-tube boxes simply tie the negative side of the HV to ground.
This amplifier doesn't. The grids of the tubes are grounded. To put any fixed bias on the tubes, the cathode can't also be grounded. As a result it "floats".
In a way.
The relay that shuts off the tubes is in the cathode circuit. The center tap of the filament transformer becomes your "return" point, as they call it. The keying relay will be connected to this center tap on one side. The other side of that relay leads back to the B-, or Bee-Minus, negative output of the HV supply. There will be a zener diode or equivalent fixed voltage-drop device in line with this relay circuit to put fixed bias onto the grids of the tubes.
The next question is what happens when the high-voltage shorts in any way to ground.
It's not shorting to its own negative side, because the Bee-minus is not grounded.
But the grid-meter shunt resistor is. That resistor will be connected to ground at one end. That's where the grids are connected, after all. The other end of the grid-meter shunt resistor goes to the Bee-Minus. This is where normal grid current flows, from grid to cathode, and the cathodes are connected to the Bee-Minus.
Whenever there is a high-voltage short to gound, the fault current will have only one path to follow. The grid-meter resistor.
And that's what brings us the the SB-220. The grid-meter resistor in this model. is at the top edge of the meter circuit board, just to the rear of the grid meter.
The usual failure mode for the factory high-voltage transformer is for the high-voltage winding inside to break down and arc to the grounded frame of the transformer. Naturally, this occurs inside the transformer's metal enclosure and won't be visible from the outside.
What you can see in the SB220 is labeled "R1" in the schemo. If the fault current from a HV short to ground goes through this part, it tends to explode into two or more pieces. And if the meter switch is set to "grid" at the time, the coil inside the right-hand meter will also vaporize.
We don't know whether your HV short to ground occurred inside the transformer or outside it. But a burned grid-meter resistor pretty well gives away that it did happen.
Thought sure I had a pic of an exploded R1 in a SB220. Wasted way too much time searching for it.
73