When you pulled the finals, did you check their numbers? Were they 2321's or 1969's?
We saw diodes on the original (they're like the MV-1Y) - a "varistor" used for bias - it was a diode but the mounting was designed for thermal contact and profiled the part it was supposed to keep in proper class of operation.
WHOAH!...(Found more attached before I could edit this...)
So, that makes me wonder, if the Bias Diodes are sucking the life out of the Base region - they are supposed to ISOLATED - usually thru a Ferrite Bead (I use a Bead and 2.2 ohm if I go back into these things) to "buffer" the voltages and RF currents from "re-rectifying" within the bias circuit can cause some transistors to "crowbar on" like a switch and blow quicker than you can turn off the power. Self-oscillation is what I'm talking about - turn the bias up too high in mA - and the parts turn on and unless loaded - which these are not - just for RF - they make very efficient switches and blow like fuses...
Place a dummy load on the Antenna jack...so the rest of this will be the test...
So when checking - you pull the test jumper that shorts all three - use TP 3 as POSITIVE (Red) and TP1 as NEGATIVE - you adjust the DRIVER bias by setting the DVM to Amps, usually 200mA or 2A - so you can trim the bias drain current - this current flows all the time the radios TX's - so it can supply the base leads a working small trickle voltage (usually about 0.65VDC) all the time it's in TX mode - this method forms it's class of operation - in this case Class AB - VR11 trims this Driver bias current - set it to about 45mA...lower than recommended is the safety margin in this radio - too high - you latch it and poof...
For the Finals, it's a little more tricky...
I' recommend that you pull those 150 ohm resistors and replace them with 180 ohm 1 watt types right now before things go any further. The 18 ohm resistors used are fine but the Diodes are shorting to ground some heavier current that reaches closer to the max forward current ratings when these are placed in SSB mode. If the 150 ohm resistors drift - they'll drift thermally - lower in resistance. This can put the ratio of 150 / 18 a bit lower and force the diodes into a runaway condition and may not continue to work properly in all modes - including FM modes - because the thermal profile talked about earlier - is affecting - stressing all parts involved. R255 and R256 are the ones I switch to 180 ohm 1 watt...
I see some things in your latest post - help me and make sure they aren't shorting out something they shouldn't - because if one finals' base shorts - the section can't get power to fire up the other final...